DsRed-Timer is a fluorescent protein published in 2000, derived from Discosoma sp.. It is reported to be a tetramer.
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DsRed-Timer was derived from DsRed with the following mutations: V105A/S197T
The mutant (E5) changes its fluorescence from green to red over time. The rate of color conversion is independent of protein concentration and therefore can be used to trace time-dependent expression.
(2000). Science, 290(5496) , 1585-1588. doi: 10.1126/science.290.5496.1585. Article
(2010). Molecular Biology of the Cell, 21(1) , 87-94. doi: 10.1091/mbc.e09-08-0722. Article
(2004). PLANT PHYSIOLOGY, 135(4) , 1879-1887. doi: 10.1104/pp.103.038539. Article